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The Diversity of Zooplankton Revealed Under the Microscope

Zooplankton is a collective term for microscopic animals that live suspended in the water column. The largest zooplankton species can just barely be seen with the naked eye, but their true diversity is revealed through a microscope. In the zooplankton image gallery, you can explore the smallest animals of the Baltic Sea!


The species in the Baltic Sea are adapted to brackish water and tend to be small compared to oceanic species. The most important zooplankton groups in our marine area are copepods (Copepoda), water fleas (Cladocera / Diplostraca), rotifers (Rotifera), and ciliates (Ciliophora).

The images in the gallery were produced as part of the Finnish Environment Institute’s zooplankton monitoring program. When using the images, the photographer’s name must be included alongside the image. The photographer’s name and additional information about the species are displayed when the image is clicked.

Copepods Are the Largest Zooplankton Group in the Baltic Sea

Copepods (Copepoda) form the largest and most important group of zooplankton in our marine area.
Copepods belong to the subclass of crustaceans (Crustacea) and undergo a complex metamorphosis during their life cycle, which includes several larval stages (nauplius larvae) and copepodite stages. The appearance of copepods changes significantly between these stages. In the final stage, a sexually mature adult emerges. Adult copepods in the Baltic Sea are about 1 mm long, with the exception of the larger Limnocalanus macrurus.

Three copepod orders are found in the Baltic Sea plankton: CalanoidaCyclopoida, and Harpacticoida. Of these, Calanoida—commonly known as calanoid copepods—are the most dominant in the Baltic Sea.

Calanoid copepods (Calanoida)

Calanoids, or calanoid copepods, are entirely planktonic species, and most of them are marine, meaning they survive only in saline water. Calanoids have an oval-shaped anterior body, and in adult individuals, the first antennae are longer than the front part of the body.

Acartia tonsa, also known as the barrel-shaped calanoid copepod, is a non-native species that arrived in the Baltic Sea in the 1920s. Acartia tonsa is highly tolerant of environmental changes and may outcompete native species of the genus Acartia if conditions change suddenly.
Acartia tonsa, male
Nauplius larvae are the first developmental stage of copepods.
Acartia sp. larva
Eurytemora affinis thrives especially in the low-salinity coastal waters throughout the Finnish marine area.
The eggs of this species are resistant to predation, meaning they do not dissolve in the digestive systems of predators.
Eurytemora affinis, female
Nauplius larvae are the first developmental stage of copepods.
Eurytemora affinis, larva
Pseudocalanus elongatus is a lover of salty water, and is therefore rarely found in the Gulf of Bothnia.
In the Baltic Sea, this species inhabits deep waters. P. elongatus is rich in fat, making it an excellent energy source for fish.
Pseudocalanus elongatus, female
Nauplius larva of the copepod Pseudocalanus elongatus.
Pseudocalanus elongatus, larva
Limnocalanus macrurus is a glacial relict and by far the largest copepod species in the Baltic Sea.
Adult individuals are 2–3 mm long. L. macrurus is a species adapted to low-salinity water, and it is found in the Gulf of Bothnia and near coastal areas. It prefers cold water, which is why it migrates to deeper waters during the summer. Due to its large size, L. macrurus is an important food source for planktivorous fish.
Limnocalanus macrurus, female
The copepod Limnocalanus macrurus undergoes several transformations before reaching adulthood.
After six nauplius larval stages, it progresses through six copepodite stages, which are juvenile phases before adulthood. The copepodites of this species are characterized by a large, distinctly curved maxilliped (feeding appendage).
Young Limnocalanus macrurus
Temora longicornis is a large surface-dwelling species. It is adapted to saline water and is therefore most abundant in the southern parts of the sea. Its presence in the Gulf of Bothnia is limited due to the low salinity of the water. Temora longicornis is an important food source for Baltic herring. Its population has declined since the 1960s.
Temora longicornis, female
Nauplius larvae are the first developmental stage of copepods.
Temora longicornis, larva
Centropages hamatus is common in the central Baltic Sea, but due to the low salinity, it is only rarely found in the Gulf of Bothnia.
Centropages hamatus, male
Nauplius larvae are the first developmental stage of copepods.
Centropages hamatus, larva

Cyclopoid copepods (Cyclopoida)

Species of the order Cyclopoida mostly live among aquatic vegetation in freshwater environments. Compared to calanoids, their anterior body is more rounded, and their first antennae are shorter. Cyclopoid copepods are primarily freshwater species.

Copepods of the order Cyclopoida, with a cyclopoid nauplius larva on the right
Cyclopoida nauplius larva

Harpacticoid copepods (Harpacticoida)

Species of the order Harpacticoida are mainly found near the sea bottom, but they are occasionally captured in plankton samples taken from the water column.

Water fleas are crustaceans

Water fleas (Cladocera/Diplostraca) belong to the crustaceans, just like copepods. Most species of water fleas inhabit freshwater environments, and among them are both herbivores and predators. Unlike copepods, water fleas develop without metamorphosis (with the exception of Leptodora kindti). Water fleas reproduce mainly asexually, through parthenogenesis, and males typically appear only in the autumn.

Two species of water fleas from the genus Bosmina can be found in the Baltic Sea: B. coregoni maritima and B. longirostris. Bosmina have a bilaterally symmetrical carapace. Their first antennae form a “snout,” which makes Bosmina easy to recognize.
Bosmina coregoni maritima
Water fleas belonging to the family Podonidae are predators, and they all have a similarly predatory eye that allows them to see widely in all directions. Species in the genus Podon are generally very similar in appearance: a rounded head and a rounded brood chamber.
Podon intermedius
Evadne anonyx also belongs to the predatory water fleas of the family Podonidae. It is a non-native (invasive) species and larger than the native E. nordmanni. In addition, E. anonyx is distinguished by a clearly visible “neck.”
Evadne anonyx
The fishhook water flea Cercopagis pengoi has spread to the Baltic Sea from the Ponto-Caspian region. This harmful invasive species has also become familiar to fishermen due to the slimy buildup it causes on fishing nets. The fishhook water flea is a predator that hunts other zooplankton using its long, hook-shaped tail.
Fishhook water flea, Cercopagis pengoi
Daphnias are hood-headed, charming water fleas. In the Baltic Sea, two main species are commonly found: D. cucullata and D. cristata, although other freshwater daphniids may occasionally appear along the coast. Daphnias feed on phytoplankton and ciliates.
Daphnia cristata
Daphnias are hood-headed, charming water fleas. In the Baltic Sea, two main species are commonly found: D. cucullata and D. cristata, although other freshwater daphniids may occasionally appear along the coast. Daphnias feed on phytoplankton and ciliates.
Daphnia cucullata
Leptodora kindtii is a large and translucent water flea, with two spines at the tip of its abdomen.
Leptodora kindtii

Rotifers (Rotifera)

Rotifers are small organisms with highly variable appearances. In the Baltic Sea, species typically range in size from about 100 to 500 µm. Rotifers get their name from a rotating, wheel-like structure at their front end, which is covered in cilia. The spinning motion of these “wheels” creates a current that directs food toward the organism’s mouth. Rotifers filter feed on dead organic matter, algae, and other microscopic living organisms in the water, making them a crucial component of the aquatic food web. The most common rotifers found in the Baltic Sea are species of Keratella and Synchaeta.

The rotifer Keratella quadrata platei is common in the Baltic Sea and is especially found near the surface in late summer.
Keratella quadrata platei
Synchaeta baltica is a very common rotifer species in the Baltic Sea.
Synchaeta baltica

Ciliates (Ciliophora)

Ciliates are single-celled protozoa less than 150 µm in length, which use cilia for movement and feeding. The cilia may cover the entire body or be concentrated in specific areas. The most common ciliates in the Baltic Sea include, for example, the genus Zoothamnium and the genera Helicostomella and Tintinnopsis, which belong to the order Tintinnoidea.

Vorticella sp. is a ciliate that feeds on small microalgae.
Vorticella sp.
Didinium sp.

Other zooplankton groups

Zooplankton samples also contain a large number of so-called meroplanktonic species, meaning species that have a planktonic stage in their life cycle. These include, for example, larval forms of certain bivalves (Bivalvia), gastropods (Gastropoda), polychaete worms (Polychaeta), and the barnacle Amphibalanus improvisus. In addition, individuals from groups such as mysid shrimps (Mysidacea) and nematodes (Nematoda) are occasionally found, as well as ephyrae, the larval stage of the moon jellyfish (Aurelia aurita). Zooplankton samples may also contain phytoplankton, fish larvae, pollen, and detritus.

The nauplius larvae of the barnacle (Amphibalanus improvisus) are only millimeters in size. In early spring, thousands of them are released into the water, after which they attach themselves headfirst to a solid surface, such as a rock, a mussel shell, or the hull of a boat.
Barnacle nauplius larva
The veliger larvae of bivalves (Bivalvia) initially live inside their parent. Once released into the water, they use tiny hooks to attach themselves to the gills or skin surface of fish, where they live parasitically for a few months before beginning an independent life.
Bivalvia veliger larva
The polychaete worm Bylgides sarsi reaches about 3 cm in length as an adult. However, during its larval stage, it is part of the zooplankton community in the Baltic Sea.
Nectochaeta larva of Bylgides sarsi